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A 62-year-old male complains to his primary care doctor about a new tremor. Exam confirms a new pill-rolling tremor. There is prominent cog-wheeling rigidity and masked facies. Which of the following most likely accounts for this patient's symptoms?

A- Neuronal degeneration of anterior horn cells
B- Neuronal degeneration of the caudate nuclei
C- Neuronal degeneration of the corticospinal tracts
D- Neuronal degeneration of spinothalamic tracts
E- Neuronal degeneration of the substantia *****
 
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